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Ricaut Francois Xavier

François-Xavier Ricaut
CNRS Researcher in Anthropobiology, University of Toulouse

Contact :
Tel : +00 33 (0)5 61 14 55 11

Research interests
Trained as an anthropobiologist, my research investigates different aspects of the history and evolution of human populations, in particular the modalities of human settlement (migration, admixture, selection, drift) that explain the genetic population structure, with a focus on the Indo-Pacific region.
This research is achieved through analysing the spatial and temporal distribution patterns of biological diversity in human populations using a wide spectrum of analytical tools. Genetic and skeletal morphological variations are investigated in both contemporary and ancient populations, allowing a fine-scale analytical perspective.

Current research
- Unexplored human history in the Indian Ocean : Human biological adaptations to the Indian Ocean slave trade and Indonesian dispersal to Madagascar
- Settlement history in Papua New Guinea : multidisciplinary approach from archaeo-sciences and molecular anthropology to reveal complex cultural and biological history
- Human occupation of karstic area, and settlement history of East Kalimantan (Indonesia)
- Genetic history of the Swahili populations
- Molecular epidemiology and population history in the Indonesian archipelago

Recently, I have begun to explore new aspects of genetic anthropology and environmental adaptation after joining the Evolutionary Medicine research group led by Dr Thierry Letellier in 2014. I have expanded my research, grounded in the anthropological framework of the Indo-Pacific and Australo-Melanesia region, to explore new scientific questions using molecular epidemiology (viral DNA and host resistance) and epigenetics. These new tools are complementary to the archaeological excavation and population genetic approaches used to reconstruct the population history of the Indo-Pacific region.

Research experience
I have been involved in 13 international projects investigating the genetic history of human populations, including human occupation in northern Asia, southern European Neolithic, Austronesian dispersal in the Indo-Pacific region, and human occupation of Sahul (Australo-Melanesia).

- 2014-2019 : Principal Investigator of the ANR (French National Research Agency) research program “OceoAdapto” (ANR-14-CE31-0013-01)
Project : Unexplored human history in the Indian Ocean
- 2016-present : Principal investigator of the French archaeological mission in Papua New Guinea.
Grant : French Ministry of Foreign Affairs
Project : Prehistory of Karawari sites, Sepik (Papua New Guinea)
- 2011-2016 : Principal investigator of the French archaeological mission in Borneo (Indonesia).
Grant : French Ministry of Foreign Affairs
Project : Human occupation of karstic area, and settlement history of East Kalimantan (Indonesia)
- 2008-present : Researcher CNRS (CR1)
Laboratoire d’Anthropologie Moléculaire et Imagerie de Synthèse (UMR5288), University of Toulouse, France
- 2008-2010 : Research Associate
Centre for Archaeological Sciences, University Catholic of Leuven, Belgium
- 2007-2008 : Post-doctoral fellow (IUAP grant)
Center for Archaeological Sciences, University Catholic of Leuven, Belgium
Project : Sagalassos Archaeological Project (Pr M Waelkens)
- 2003-2006 : Post-doctoral fellow (NERC grant)
Leverhulme Centre for Human Evolutionary Studies (LCHES), Department of Biological Anthropology, University of Cambridge, UK
Project : Pioneers of Island Melanesia (Pr R Foley and Dr M Mirazon-Lahr)
- 2000-2003 : Ph.D. in Molecular Anthropology-Archaeology
Ecole des Hautes Etudes en Science Sociales and University of Toulouse (France)
Project : Ancient DNA and Human populations : Settlement of Oriental Siberia and Altaï (Pr E Crubézy and Pr B Ludes)

52 peer-reviewed publications.9 book chapters, 46 conferences.

Details available at

- Kusuma P, Brucato N, Cox MP, Letellier T, Manan A, Nuraini C, Grangé P, Sudoyo H, RICAUT FX. 2017. The last sea nomads of the Indonesian archipelago : genomic origins and dispersal. Eur J Hum Genet. 2017 Aug ;25(8):1004-1010 doi : 10.1038/ejhg.2017.88 PDF

- Brucato N, Kusum P, Beaujard P, Sudoyo H, Cox MP, RICAUT FX. Genomic admixture tracks pulses of economic activity over 2,000 years in the Indian Ocean trading network. Scientific Report 7:2919 doi : 10.1038/s41598-017-03204-y. PDF

- Pierron D, Heiske M, Razafindrazaka H, Rakoto I, Rabetokotany N, Ravololomanga B, Rakotozafy LM, Rakotomalala MM, Razafiarivony M, Rasoarifetra B, Raharijesy MA, Razafindralambo L, Ramilisonina, Fanony F, Lejamble S, Thomas O, Mohamed Abdallah A, Rocher C, Arachiche A, Tonaso L, Pereda-Loth V, Schiavinato S, Brucato N, RICAUT FX, Kusuma P, Sudoyo H, Ni S, Boland A, Deleuze JF, Beaujard P, Grange P, Adelaar S, Stoneking M, Rakotoarisoa JA, Radimilahy C, Letellier T. Genomic landscape of human diversity across Madagascar. Proc Natl Acad Sci U S A. 2017 Aug 8 ;114(32):E6498-E6506. doi : 10.1073/pnas.1704906114. PDF

- Inchley CE, Larbey CD, Shwan NA, Pagani L, Saag L, Antão T, Jacobs G, Hudjashov G, Metspalu E, Mitt M, Eichstaedt CA, Malyarchuk B, Derenko M, Wee J, Abdullah S, RICAUT FX, Mormina M, Mägi R, Villems R, Metspalu M, Jones MK, Armour JA, Kivisild T. Selective sweep on human amylase genes postdates the split with Neanderthals. Sci Rep. 2016 Nov 17 ;6:37198. doi:10.1038/srep37198. PDF

- Skoglund P, Posth C, Sirak K, Spriggs M, Valentin F, Bedford S, Clark GR, Reepmeyer C, Petchey F, Fernandes D, Fu Q, Harney E, Lipson M, Mallick S, Novak M, Rohland N, Stewardson K, Abdullah S, Cox MP, Friedlaender FR, Friedlaender JS, Kivisild T, Koki G, Kusuma P, Merriwether DA, RICAUT FX, Wee JT, Patterson N, Krause J, Pinhasi R, Reich D. Genomic insights into the peopling of the Southwest Pacific. Nature. 2016 Oct 3 ;538(7626):510-513. doi : 10.1038/nature19844. PDF

- Pagani L, Lawson DJ, Jagoda E, Mörseburg A, Eriksson A, Mitt M, Clemente F, Hudjashov G, DeGiorgio M, Saag L, Wall JD, Cardona A, Mägi R, Sayres MA, Kaewert S, Inchley C, Scheib CL, Järve M, Karmin M, Jacobs GS, Antao T, Iliescu FM, Kushniarevich A, Ayub Q, Tyler-Smith C, Xue Y, Yunusbayev B, Tambets K, Mallick CB, Saag L, Pocheshkhova E, Andriadze G, Muller C, Westaway MC, Lambert DM, Zoraqi G, Turdikulova S, Dalimova D, Sabitov Z, Sultana GN, Lachance J, Tishkoff S, Momynaliev K, Isakova J, Damba LD, Gubina M, Nymadawa P, Evseeva I, Atramentova L, Utevska O, RICAUT FX, Brucato N, Sudoyo H, Letellier T, Cox MP, Barashkov NA, Škaro V, Mulahasanovic L, Primorac D, Sahakyan H, Mormina M, Eichstaedt CA, Lichman DV, Abdullah S, Chaubey G, Wee JT, Mihailov E, Karunas A, Litvinov S, Khusainova R, Ekomasova N, Akhmetova V, Khidiyatova I, Marjanović D, Yepiskoposyan L, Behar DM, Balanovska E, Metspalu A, Derenko M, Malyarchuk B, Voevoda M, Fedorova SA, Osipova LP, Lahr MM, Gerbault P, Leavesley M, Migliano AB, Petraglia M, Balanovsky O, Khusnutdinova EK, Metspalu E, Thomas MG, Manica A, Nielsen R, Villems R, Willerslev E, Kivisild T, Metspalu M. Genomic analyses inform on migration events during the peopling of Eurasia. Nature. 2016 Oct 13 ;538(7624):238-242. doi : 10.1038/nature19792. PDF

- Malaspinas AS, Westaway MC, Muller C, Sousa VC, Lao O, Alves I, Bergström A, Athanasiadis G, Cheng JY, Crawford JE, Heupink TH, Macholdt E, Peischl S, Rasmussen S, Schiffels S, Subramanian S, Wright JL, Albrechtsen A, Barbieri C, Dupanloup I, Eriksson A, Margaryan A, Moltke I, Pugach I, Korneliussen TS, Levkivskyi IP, Moreno-Mayar JV, Ni S, Racimo F, Sikora M, Xue Y, Aghakhanian FA, Brucato N, Brunak S, Campos PF, Clark W, Ellingvåg S, Fourmile G, Gerbault P, Injie D, Koki G, Leavesley M, Logan B, Lynch A, Matisoo-Smith EA, McAllister PJ, Mentzer AJ, Metspalu M, Migliano AB, Murgha L, Phipps ME, Pomat W, Reynolds D, RICAUT FX, Siba P, Thomas MG, Wales T, Wall CM, Oppenheimer SJ, Tyler-Smith C, Durbin R, Dortch J, Manica A, Schierup MH, Foley RA, Lahr MM, Bowern C, Wall JD, Mailund T, Stoneking M, Nielsen R, Sandhu MS, Excoffier L, Lambert DM, Willerslev E.A genomic history of Aboriginal Australia. Nature. 2016 Oct 13 ;538(7624):207-214. doi : 10.1038/nature18299. PDF

- Brucato N, Kusuma P, Cox MP, Pierron D, Purnomo GA, Adelaar A, Kivisild T, Letellier T, Sudoyo H, RICAUT FX. Malagasy Genetic Ancestry Comes from an Historical Malay Trading Post in Southeast Borneo. Mol Biol Evol. 2016 Sep ;33(9):2396-400. doi : 10.1093/molbev/msw117. PDF

- Mörseburg A, Pagani L, RICAUT FX, Yngvadottir B, Harney E, Castillo C, Antao T, Hoogervorst T, Kusuma P, Razafindrazaka H,Cardona A, Pierron D, Letellier T, Wee J, Abdullah MS, Metspalu M, Kivisild T. 2015. A genomic history of Aboriginal Australia. Multi-layered population structure and local adaptations in Island Southeast Asians. Eur J Hum Genet 2016 Nov ;24(11):1605-1611 doi:10.1038/ejhg.2016.60 PDF

- Grenet M., Sarel J., Fauzy R., Oktaviana A.A., Sugiyanto B., Chazine J.-M., RICAUT FX. “New insights on the late Pleistocene-Holocene lithic industry in East Kalimantan (Borneo). The contribution of three rock shelter sites in the karstic area of the Mangkalihat peninsula”. Quaternary International (2016),416, 126-150. PDF

- Kusuma P, Brucato N, Cox MP, Pierron D, Razafindrazaka H, Sudoyo H, Letellier T, FX RICAUT. 2016. Contrasting Linguistic and Genetic Influences during the Austronesian Settlement of Madagascar. Sci Rep.6, 26066 ; doi : 10.1038/srep26066 PDF

- Plutniak S., Astolfo A., Puaud S., Ferrié J.-G., Oktaviana A.A., Sugiyanto B., Chazine J.-M., RICAUT FX. “Borneo as a half emptypot : pottery assemblage from Liang Abu, East Kalimantan”. Quaternary International 416, 228-242. doi:10.1016/j.quaint.2015.11.080 PDF

- Raghavan M, Steinrücken M, Harris K, Schiffels S, Rasmussen S, DeGiorgio M, Albrechtsen A, Valdiosera C, Ávila-Arcos MC, Malaspinas AS, Eriksson A, Moltke I, Metspalu M, Homburger JR, Wall J, Cornejo OE, Moreno-Mayar JV, Korneliussen TS, Pierre T, Rasmussen M, Campos PF, Damgaard Pde B, Allentoft ME, Lindo J, Metspalu E, Rodríguez-Varela R, Mansilla J, Henrickson C, Seguin-Orlando A, Malmström H, Stafford T Jr, Shringarpure SS, Moreno-Estrada A, Karmin M, Tambets K, Bergström A, Xue Y, Warmuth V, Friend AD, Singarayer J, Valdes P, Balloux F, Leboreiro I, Vera JL, Rangel-Villalobos H, Pettener D, Luiselli D, Davis LG, Heyer E, Zollikofer CP, Ponce de León MS, Smith CI, Grimes V, Pike KA, Deal M, Fuller BT, Arriaza B, Standen V, Luz MF, RICAUT FX., Guidon N, Osipova L, Voevoda MI, Posukh OL, Balanovsky O, Lavryashina M, Bogunov Y, Khusnutdinova E, Gubina M, Balanovska E, Fedorova S, Litvinov S, Malyarchuk B, Derenko M, Mosher MJ, Archer D, Cybulski J, Petzelt B, Mitchell J, Worl R, Norman PJ, Parham P, Kemp BM, Kivisild T, Tyler-Smith C, Sandhu MS, Crawford M, Villems R, Smith DG, Waters MR, Goebel T, Johnson JR, Malhi RS, Jakobsson M, Meltzer DJ, Manica A, Durbin R, Bustamante CD, Song YS, Nielsen R, Willerslev E. 2015 Genomic evidence for the Pleistocene and recent population history of Native Americans. Science. 2015 Aug 21 ;349(6250):aab3884 PDF

- Kusuma P, Cox MP, Pierron D, Razafindrazaka H, Brucato N, Adelaar S, Letellier T, Sudoyo H, FX RICAUT. 2015. MitochondrialDNA and the Y chromosome suggest the settlement of Madagascar by Indonesian sea nomad populations. BMC Genomics17 ;16:191 PDF

- Karmin M, Saag L, Vicente M, Wilson Sayres MA, Järve M, Talas UG, Rootsi S, Ilumäe AM, Mägi R, Mitt M, Pagani L, Puurand T, Faltyskova Z, Clemente F, Cardona A, Metspalu E, Sahakyan H, Yunusbayev B, Hudjashov G, DeGiorgio M, Loogväli EL, Eichstaedt C, Eelmets M, Chaubey G, Tambets K, Litvinov S, Mormina M, Xue Y, Ayub Q, Zoraqi G, Korneliussen TS, Akhatova F, Lachance J, Tishkoff S, Momynaliev K, FX RICAUT, Kusuma P, Razafindrazaka H, Pierron D, Cox MP, Sultana GN, Willerslev R, Muller C, Westaway M, Lambert D, Skaro V, Kovačevic L, Turdikulova S, Dalimova D, Khusainova R, Trofimova N, Akhmetova V, Khidiyatova I, Lichman DV, Isakova J, Pocheshkhova E, Sabitov Z, Barashkov NA, Nymadawa P, Mihailov E, Seng JW, Evseeva I, Migliano AB, Abdullah S, Andriadze G, Primorac D, Atramentova L, Utevska O, Yepiskoposyan L, Marjanovic D, Kushniarevich A, Behar DM, Gilissen C, Vissers L, Veltman JA, Balanovska E, Derenko M, Malyarchuk B, Metspalu A, Fedorova S, Eriksson A, Manica A, Mendez FL, Karafet TM, Veeramah KR, Bradman N, Hammer MF, Osipova LP, Balanovsky O, Khusnutdinova EK, Johnsen K, Remm M, Thomas MG, Tyler-Smith C, Underhill PA, Willerslev E, Nielsen R, Metspalu M, Villems R, Kivisild T. 2015. A recent bottleneck of Y chromosome diversity coincides witha global change in culture. Genome Res. 25(4):459-66. PDF

- Kusuma P, Cox M. P., Brucato N., Sudoyo H., Letellier T., FX RICAUT. 2015. A Western Eurasian genetic influences in the Indonesian Archipelago. Quaternary International. 416, 243-248. doi : 10.1038/srep26066. PDF

- Pierron D, H Razafindrazaka, L Pagani, FX RICAUT, T Antao, M Capredon, C Sambo, C Radimilahy, JA Rakotoarisoa, R Blench,T Letellier, T Kivisild. 2014. Genome-wide evidence of Austronesian-Bantu admixture and cultural reversion in a huntergatherergroup of Madagascar.Proc Natl AcadSci U S A. 111, p. 936–941 PDF

- Plutniak, S., A. A. Oktaviana, J.-G. Ferrié, B. Sugiyanto, J. Sarels, A. Guerreiro, R. Fauzi, M. Grenet, T. Simanjuntak et RICAUT FX. 2014. Human Occupation of the Mangkalihat Karstic Area : Rock Art and Settlement History in EastKalimantan. Borneo Research Bulletin, 45, p. 320–322.

- Plutniak S, Oktaviana AA, Sugiyanto B, Grenet M, J.-M. Chazine, RICAUT FX.. 2013. New ceramic data from East Kalimantan : Pottery chronology and the cord-marked and red-slipped sherds of Liang Abu’s layer 2. Journal of Pacific Archaeology vol. 5, no 1, p.90-99. PDF